Most of these articles are available for download from Research Gate at the following link.

*student authors from Eastern and UNLV


Dolby, G.A., R.J. Dorsey & M.R. Graham. Charting the biogeographic legacy of the lower Colorado River: new insights from geology and comparative phylogeography of 28 desert fauna. In review.

Graham, M.R., M.B. Pinto*, & P.E. Cushing.  A quick test of the light attraction hypothesis in camel spiders of the Mojave Desert (Arachnida: Solifugae). In review.

Bryson, R.W., D.A. Wood, M.R. Graham, M.E. Soleglad & J.E. McCormack. 2018. Species delimitation using genome-wide SNP data supports the distinction of two new species of scorpions endemic to California (Arachnida: Scorpiones: Kovarikia). ZooKeys, 739: 79–106. [new species: Kovarikia savaryi and K. oxy]

Fet, V., F. Kovarik, B. Gantenbein, R.C. Kaiser*, A.K. Stewart & M.R. Graham. 2018. Revision of the Mesobuthus caucasicus complex from Central Asia, with descriptions of six new species (Scorpiones: Buthidae). Euscorpius, 255:1–77. [new species: Mesobuthus brutus, M. elenae, M. gorelovi, M. kreuzbergi, M. mischis, and M. nenilini]


Román, J.P., F.C. García, D. Medina, M. Vásquez, J. García, M.R. Graham, D. Romero-Álvarez, P.O. Pardal, E.A.Y. Ishikawa & A. Borges. 2017. Scorpion envenoming in Morona Santiago, Amazonian Ecuador: Molecular phylogenetics confirms the involvement of the Tityus obscurus group. Acta Tropica, 178:1–9.

Soleglad, M.E., R.F. Ayrey, M.R. Graham & V. Fet.  2017. Catalinia, a new scorpion genus from southern California, USA and northern Baja California, Mexico (Scorpiones: Vaejovidae). Euscorpius, 251:1–64. [new genus: Catalinia]

Graham, M.R., D.A. Wood., J. Henault*, Z.J. Valois* & P.E. Cushing. 2017. Ancient lakes, Pleistocene climates, and river avulsions structure the phylogeography of a large but little-known rock scorpion from the Mojave and Sonoran deserts. Biological Journal of the Linnean Society, 122(1):133­–146.


Soleglad, M.E., V. Fet, M.R. Graham & R.F.Ayrey. 2016. Graemeloweus, a new scorpion genus from northern California, USA (Scorpiones: Vaejovidae). Euscorpius, 227:1–38. [new genus: Graemeloweus]

Sissom, W.D., M.R. Graham, T.G. Donaldson & R.W. Bryson. 2016. Two new Vaejovis C.L. Koch 1836 from highlands of the Sierra Madre Occidental, Durango, Mexico (Scorpiones, Vaejovidae). Insecta Mundi, 477:1–14. [new species: Vaejovis sierrae, V. mcwesti]

Fet, V., M.R. Graham, G. Blagoev, A. Karataş & Ah. Karataş. 2016. DNA barcoding indicates hidden diversity of Euscorpius (Scorpiones: Euscorpiidae) in Turkey. Euscorpius, 216:1–12.


Cushing, P. E., M.R. Graham, L. Prendini & J.O. Brookhart. 2015. A multilocus molecular phylogeny of the endemic North American camel spider family Eremobatidae (Arachnida: Solifugae). Molecular Phylogenetics and Evolution, 92:280–293.

Graham, M.R., B.E. Hendrixson, C.A. Hamilton & J.E. Bond. 2015. Miocene extensional tectonics explain ancient patterns of diversification among turret-building tarantulas (Aphonopelma mojave group) in the Mojave and Sonoran deserts. Journal of Biogeography, 42(6):1052–1065.


Graham, M.R., R.W. Bryson & B.R. Riddle. 2014. Late Pleistocene to Holocene distributional stasis in scorpions along the Baja California Peninsula. Biological Journal of the Linnean Society, 11:450–461.

Borges, A. & M.R. Graham. 2014. Phylogenetics of Scorpions of Medical Importance. Pp. 1-20, in Toxinology. Venom Genomics and Proteomics (P. Gopalakrishnakone, ed.). Springer. ISBN: 978-94-007-6649-5

Fet, V., M.R. Graham, M.M. Webber & G. Blagoev. 2014. Two new species of Euscorpius (Scorpiones: Euscorpiidae) from Bulgaria, Serbia, and Greece. Zootaxa, 3894:083–105. [new species: Euscorpius deltshevi, E. solegladi]

Soleglad, M.E., V. Fet & M.R. Graham. 2014. Kovarikia, a new scorpion genus from California, USA (Scorpiones: Vaejovidae). Euscorpius, 185:1–22. [new genus: Kovarikia]


Graham, M.R., J.R. Jaeger, L. Prendini & B.R. Riddle. 2013. Phylogeography of Beck’s Desert Scorpion, Paruroctonus becki, reveals Pliocene diversification in the Eastern California Shear Zone and postglacial expansion in the Great Basin Desert. Molecular Phylogenetics and Evolution, 69:502–513.

Webber, M.M. & M.R. Graham. 2013. An Arizona bark scorpion (Centruroides sculpturatus) found consuming a venomous prey item nearly twice its length. Western North American Naturalist, 73:530–532.

Graham, M.R., J.R. Jaeger, L. Prendini & B.R. Riddle. 2013. Phylogeography of the Arizona hairy scorpion (Hadrurus arizonensis) supports a model of biotic assembly in the Mojave Desert and adds a new Pleistocene refugium. Journal of Biogeography, 40:1298–1312.

Tate, A.E.*, R.R. Riddle*, M.E. Soleglad & M.R. Graham. 2013. Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City,” Nevada (Scorpiones, Vaejovidae). ZooKeys, 364:29–45. [new species: Pseudouroctonus peccatum]

Bryson, R.W., B.R. Riddle, M.R. Graham, B.T. Smith & L. Prendini. 2013. As old as the hills: montane scorpions in southwestern North America reveal an ancient association between biotic diversification and landscape history. PLoS ONE 8:e52822.


Graham, M.R., M.M. Webber, G. Blagoev, N. Ivanova & V. Fet. 2012. Molecular and morphological evidence support the elevation of Euscorpius germanus croaticus Di Caporiacco, 1950 (Scorpiones: Euscorpiidae) to E. croaticus stat. nov., a rare species from Croatia. Revista Ibérica de Aracnologia, 21:41–50. [new status: Euscorpius croaticus]

Graham, M.R., R.F. Ayrey & R.B. Bryson. 2012. Multivariate methods support the distinction of a new highland Vaejovis (Scorpiones: Vaejovidae) from the Sierra de los Ajos, Mexico. Journal of Arachnology, 40(3):281–290. [new species: Vaejovis bandido]

Graham, M.R., P. Stoev, N. Akkari, G. Blagoev & V. Fet. 2012. Euscorpius sicanus (Scorpiones: Euscorpiidae) from Tunisia: DNA barcoding confirms ancient disjunctions across the Mediterranean Sea. Serket, 13(1/2):16–26.

Webber, M.M., M.R. Graham & J.R. Jaeger. 2012. Wernerius inyoensis, an elusive new scorpion from the Inyo Mountains of California (Scorpiones: Vaejovidae). ZooKeys, 177:1–13. [new species: Wernerius inyoensis; article featured by MSNBC, LA Times, Chicago Tribune, Our Amazing Planet, Eureka Alert, and numerous other international media outlets]

Graham, M.R., V. Oláh-Hemmings & V. Fet. 2012. Phylogeography of co-distributed dune scorpions identifies the Amu Darya River as a long-standing component of Central Asian biogeography. Zoology in the Middle East, 55:95–110.


Bryson, R.B, R.W. Murphy, M.R. Graham, A. Lathrop & D. Lazcano-villarea. 2011. Ephemeral Pleistocene woodlands connect the dots for highland rattlesnakes of the Crotalus intermedius group. Journal of Biogeography, 38:2299–2310.

Graham, M.R. 2011. Book Review – Scorpions of the World. Journal of Arachnology, 39:166–167.

Crother, B.I., M.E. White, J.M. Savage, M.E. Eckstut, M.R. Graham & D.W. Gardner. 2011. A Reevaluation of the Status of the Foxsnakes Pantherophis gloydi Conant and P. vulpinus Baird and Girard (Lepidosauria). ISRN Zoology 1–15. [new species: Pantherophis ramspotti]


Graham, M.R. & R.W. Bryson. 2010. Vaejovis montanus (Scorpiones: Vaejovidae), a new species from the Sierra Madre Occidental of Mexico. Journal of Arachnology, 38:285–293.

Bryson, R.W. & M.R. Graham. 2010. A new alligator lizard from northeastern Mexico. Herpetologica, 66:92–98.


Graham, M.R. & M.E. Soleglad. 2007. A new scorpion genus representing a primitive taxon of tribe Stahnkeini, with a description of a new species from Sonora, Mexico(Scorpiones: Vaejovidae). Euscorpius, 57:1–13. [new genus & species: Gertschius crassicorpus]

Graham, M.R. 2007. Sky Island Vaejovis: two new species and a redescription of V. vorhiesi Stahnke (Scorpiones: Vaejovidae). Euscorpius, 51:1–14. [new species: Vaejovis cashi, V. feti]


Teruel, R., V. Fet & M.R. Graham. 2006. The first mitochondrial DNA phylogeny of Cuban Buthidae (Scorpiones: Buthoidea). Boletín de la S.E.A., 39:219–226.

Graham, M.R. & V. Fet. 2006. Serrula in retrospect: a historical look at scorpion literature (Scorpiones: Orthosterni). Euscorpius, 48:1–19.

Graham, M.R. 2006. Redescription and lectotype designation of Vaejovis lapidicola Stahnke, 1940 (Scorpiones: Vaejovidae). Euscorpius, 46:1–6.

Graham, M.R. 2006. Malformed pedipalp finger dentition of the scorpion Superstitionia donensis (Scorpiones: Superstitioniidae). Euscorpius, 42:1–4.


Fan, J., M.R. Graham, H. Akabane, L.L. Richardson & G. Zhu. 2005. Identification of a novel male germ cell-specific gene TESF-1 in mice. Biochemical and Biophysical Research Communications, 340:8–12.


Fet, E.V., D. Neff, M.R. Graham & V. Fet. 2003. Metasoma of Orthochirus (Scorpiones: Buthidae): are scorpions evolving a new sensory organ? Revista Ibérica de Aracnologia, 8:69–72.