PUBLICATIONS

Most of these articles are available for download from Research Gate at the following link.

https://www.researchgate.net/profile/Matthew_Graham2

*Student authors

Kulkarni, S., Steiner, H., Garcia, E., Iuri, H., Jones, R., Ballesteros, J., … & Sharma, P. P. (2022). Neglected no longer: Phylogenomic resolution of higher-level relationships in Solifugae. bioRxiv, 2022-10. Preprint. 

2023

Santibáñez-López, C.E., A.A. Ojanguren-Affilastro, M.R. Graham & P.P. Sharma. (Accepted) Congruence between ultraconserved element-based matrices and phylotranscriptomic datasets in the scorpion tree of life. Cladistics.

Atkinson*, L., C. Shimwell, K.M. Lucin, M.R. Graham & B. Murdoch. (In Press). Antibiotic-producing bacteria isolated from the giant sand scorpion, Smeringurus mesaensis (Scorpiones: Vaejovidae). Journal of Arachnology.

Shimwell*, C., L. Atkinson*, M.R. Graham & B. Murdoch. (2023). A first molecular characterization of the scorpion telson microbiota of Hadrurus arizonensis and Smeringurus mesaensis. PloS ONE, 18(1), e0277303.

2022

Poverennyi, N.M., M.R. Graham & V.Y. Fet. 2022. The northernmost Palearctic scorpion. Arthropoda Selecta, 32(2): 2013–2016.

Kovařík, F., V. Fet, B, Gantenbein, M.R. Graham, E.A. Yağmur, F. Šťáhlavský, N.M. Poverennyi & N.E. Novruzov. 2022. A revision of genus Mesobuthus Vachon, 1950, with a description of 14 new species (Scorpiones: Buthidae). Euscorpius, 348: 1–189. [new species: Mesobuthus birulai, M. crucittii, M. farleyi, M. fomichevi, M. galinae, M. kaftani, M. marusiki, M. mirshamsii, M. navidpouri, M. rahsenae, M. turcicus, M. vignolii, M. yagmuri, and M. zonsteini]

2021

Santibañez-López, C.E., P.E. Cushing, A.M. Powell* & M.R. Graham. 2021. Diversification, introgression, and post-glacial range expansion of giant North American camel spiders in genus Eremocosta (Solifugae: Eremobatidae). Scientific Reports, 11: 22093.

Borges, A., M.R. Graham, D.M. Candido & P.P. Oliveira Pardal. 2021. Amazonian scorpions and scorpionism: integrating toxinological, clinical, and phylogenetic data to combat a human health crises in the world’s most diverse rainforest. Journal of Venomous Animals and Toxins including Tropical Diseases, 27: e2021002.

Graham, M.R., E.L. Garcia*, B.E. Hendrixson, A.M. Sampognaro* & P.E. Cushing. 2021. Pliocene origins, Pleistocene refugia, and postglacial range expansions in southern devil scorpions (Vaejovidae: Vaejovis carolinianus). Organisms Diversity & Evolution, 21(3): 575–590.

Fet, V., F. Kovařík, B. Gantenbein & M.R. Graham. 2021. Three new species of Olivierus (Scorpiones: Buthidae) from Kazakhstan and Uzbekistan. Zootaxa, 5006(1): 54–72. https://doi.org/10.11646/zootaxa.5006.1.9 [new species: Olivierus mikhailovi, O. tarabaevi, and O. voldemari]

Azzinnari, J.S.*, R.W. Bryson Jr., M.R. Graham, C. Solís-Rojas & W.D. Sissom. 2021. A new Vaejovis C.L. Koch from the Sierra Madre Occidental of Durango, Mexico (Scorpiones: Vaejovidae). Insecta Mundi 0852: 1–12. [new species: Vaejovis baggins]

Santibáñez-López, C.E, K. Farleigh*, P.E. Cushing & M.R. Graham. 2021. Restriction enzyme optimization for RADseq with camel spiders (Arachnida: Solifugae). Journal of Arachnology 8: 346–350.

2020

Borges, A., A.R. de Arias, S. de Almeida Lima, B. Lomonte, C. Díaz, C. Chávez-Olórtegui, M.R. Graham, E. Kalapothakis, C. Coronel & A.R. de Roodt. 2020. Genetic and toxinological divergence among populations of Tityus trivittatus Kraepelin, 1898 (Scorpiones: Buthidae) inhabiting Paraguay and Argentina. PLoS Neglected Tropical Diseases 14(12): e0008899.

Graham, M.R., C.E. Santibáñez-López, S. Derkarabetian & B.E. Hendrixson. 2020. Pleistocene persistence and expansion in tarantulas on the Colorado Plateau and the effects of missing data on phylogeographical inferences from RADseq. Molecular Ecology 29(19): 3684–3701.

Borges, A., B. Lomonte, Y. Angulo, H.A. de Patiño, J.M. Pascale, R. Otero, R.J. Miranda, L. De Sousa, M.R. Graham, A. Gómez, P.P.O. Pardal, E. Ishikawa, F. Bonilla, A. Castillo, R.A.M. de Avila, J.P. Gómez & J.A. Caro-López. 2020. Venom diversity in the Neotropical scorpion genus Tityus: implications for antivenom design emerging from molecular and immunochemical analyses across endemic areas of scorpionism. Acta Tropica 204: 105346

2019

Santibáñez-López, C.E., M.R. Graham, P.P. Sharma, E. Ortiz & L.D. Possani. 2019. Hadrurid Scorpion Toxins: Evolutionary Conservation and Selective Pressures. Toxins 11(11): 637.

Graham, M.R., E.D. Myers, R.C. Kaiser* & V. Fet. 2019. Cryptic species and co-diversification in sand scorpions from the Karakum and Kyzylkum Deserts of Central Asia. Zoologica Scripta 48(6): 801–812.

Dolby, G.A., R.J. Dorsey & M.R. Graham. 2019. A legacy of geo-climatic complexity and genetic divergence along the lower Colorado River: Insights from the geologic record and 33 desert-adapted animals. Journal of Biogeography 46(11): 2479–2505.

Graham, M.R., M.B. Pinto* & P.E. Cushing. 2019. A test of the light attraction hypothesis in camel spiders of the Mojave Desert (Arachnida: Solifugae). Journal of Arachnology 47(2): 293–296.

2018

Bryson, R.W., D.A. Wood, M.R. Graham, M.E. Soleglad & J.E. McCormack. 2018. Species delimitation using genome-wide SNP data supports the distinction of two new species of scorpions endemic to California (Arachnida: Scorpiones: Kovarikia). ZooKeys, 739: 79–106. [new species: Kovarikia savaryi and K. oxy]

Fet, V., F. Kovarik, B. Gantenbein, R.C. Kaiser*, A.K. Stewart & M.R. Graham. 2018. Revision of the Mesobuthus caucasicus complex from Central Asia, with descriptions of six new species (Scorpiones: Buthidae). Euscorpius 255: 1–77. [new species: Mesobuthus brutus, M. elenae, M. gorelovi, M. kreuzbergi, M. mischis, and M. nenilini]

2017

Román, J.P., F.C. García, D. Medina, M. Vásquez, J. García, M.R. Graham, D. Romero-Álvarez, P.O. Pardal, E.A.Y. Ishikawa & A. Borges. 2017. Scorpion envenoming in Morona Santiago, Amazonian Ecuador: Molecular phylogenetics confirms the involvement of the Tityus obscurus group. Acta Tropica 178: 1–9.

Soleglad, M.E., R.F. Ayrey, M.R. Graham & V. Fet.  2017. Catalinia, a new scorpion genus from southern California, USA and northern Baja California, Mexico (Scorpiones: Vaejovidae). Euscorpius 251: 1–64. [new genus: Catalinia]

Graham, M.R., D.A. Wood., J. Henault*, Z.J. Valois* & P.E. Cushing. 2017. Ancient lakes, Pleistocene climates, and river avulsions structure the phylogeography of a large but little-known rock scorpion from the Mojave and Sonoran deserts. Biological Journal of the Linnean Society 122(1): 133­–146.

2016

Soleglad, M.E., V. Fet, M.R. Graham & R.F.Ayrey. 2016. Graemeloweus, a new scorpion genus from northern California, USA (Scorpiones: Vaejovidae). Euscorpius 227: 1–38. [new genus: Graemeloweus]

Sissom, W.D., M.R. Graham, T.G. Donaldson & R.W. Bryson. 2016. Two new Vaejovis C.L. Koch 1836 from highlands of the Sierra Madre Occidental, Durango, Mexico (Scorpiones, Vaejovidae). Insecta Mundi 477: 1–14. [new species: Vaejovis sierrae, V. mcwesti]

Fet, V., M.R. Graham, G. Blagoev, A. Karataş & Ah. Karataş. 2016. DNA barcoding indicates hidden diversity of Euscorpius (Scorpiones: Euscorpiidae) in Turkey. Euscorpius 216: 1–12.

2015

Cushing, P. E., M.R. Graham, L. Prendini & J.O. Brookhart. 2015. A multilocus molecular phylogeny of the endemic North American camel spider family Eremobatidae (Arachnida: Solifugae). Molecular Phylogenetics and Evolution 92: 280–293.

Graham, M.R., B.E. Hendrixson, C.A. Hamilton & J.E. Bond. 2015. Miocene extensional tectonics explain ancient patterns of diversification among turret-building tarantulas (Aphonopelma mojave group) in the Mojave and Sonoran deserts. Journal of Biogeography 42(6): 1052–1065.

2014

Graham, M.R., R.W. Bryson & B.R. Riddle. 2014. Late Pleistocene to Holocene distributional stasis in scorpions along the Baja California Peninsula. Biological Journal of the Linnean Society 11(2): 450–461.

Borges, A. & M.R. Graham. 2014. Phylogenetics of Scorpions of Medical Importance. Pp. 1-20, in Toxinology. Venom Genomics and Proteomics (P. Gopalakrishnakone, ed.). Springer. ISBN: 978-94-007-6649-5

Fet, V., M.R. Graham, M.M. Webber & G. Blagoev. 2014. Two new species of Euscorpius (Scorpiones: Euscorpiidae) from Bulgaria, Serbia, and Greece. Zootaxa 3894(1): 083–105. [new species: Euscorpius deltshevi, E. solegladi]

Soleglad, M.E., V. Fet & M.R. Graham. 2014. Kovarikia, a new scorpion genus from California, USA (Scorpiones: Vaejovidae). Euscorpius 185:1 –22. [new genus: Kovarikia]

2013

Graham, M.R., J.R. Jaeger, L. Prendini & B.R. Riddle. 2013. Phylogeography of Beck’s Desert Scorpion, Paruroctonus becki, reveals Pliocene diversification in the Eastern California Shear Zone and postglacial expansion in the Great Basin Desert. Molecular Phylogenetics and Evolution 69(3): 502–513.

Webber, M.M. & M.R. Graham. 2013. An Arizona bark scorpion (Centruroides sculpturatus) found consuming a venomous prey item nearly twice its length. Western North American Naturalist 73(4): 530–532.

Graham, M.R., J.R. Jaeger, L. Prendini & B.R. Riddle. 2013. Phylogeography of the Arizona hairy scorpion (Hadrurus arizonensis) supports a model of biotic assembly in the Mojave Desert and adds a new Pleistocene refugium. Journal of Biogeography 40(7): 1298–1312.

Tate, A.E.*, R.R. Riddle*, M.E. Soleglad & M.R. Graham. 2013. Pseudouroctonus peccatum, a new scorpion from the Spring Mountains near “Sin City,” Nevada (Scorpiones, Vaejovidae). ZooKeys 364: 29–45. [new species: Pseudouroctonus peccatum]

Bryson, R.W., B.R. Riddle, M.R. Graham, B.T. Smith & L. Prendini. 2013. As old as the hills: montane scorpions in southwestern North America reveal an ancient association between biotic diversification and landscape history. PLoS ONE 8: e52822.

2012

Graham, M.R., M.M. Webber, G. Blagoev, N. Ivanova & V. Fet. 2012. Molecular and morphological evidence support the elevation of Euscorpius germanus croaticus Di Caporiacco, 1950 (Scorpiones: Euscorpiidae) to E. croaticus stat. nov., a rare species from Croatia. Revista Ibérica de Aracnologia, 21: 41–50. [new status: Euscorpius croaticus]

Graham, M.R., R.F. Ayrey & R.B. Bryson. 2012. Multivariate methods support the distinction of a new highland Vaejovis (Scorpiones: Vaejovidae) from the Sierra de los Ajos, Mexico. Journal of Arachnology, 40(3):281–290. [new species: Vaejovis bandido]

Graham, M.R., P. Stoev, N. Akkari, G. Blagoev & V. Fet. 2012. Euscorpius sicanus (Scorpiones: Euscorpiidae) from Tunisia: DNA barcoding confirms ancient disjunctions across the Mediterranean Sea. Serket, 13(1/2):16–26.

Webber, M.M., M.R. Graham& J.R. Jaeger. 2012. Wernerius inyoensis, an elusive new scorpion from the Inyo Mountains of California (Scorpiones: Vaejovidae). ZooKeys, 177:1–13. [new species: Wernerius inyoensis; article featured by MSNBC, LA Times, Chicago Tribune, Our Amazing Planet, Eureka Alert, and numerous other international media outlets]

Graham, M.R., V. Oláh-Hemmings & V. Fet. 2012. Phylogeography of co-distributed dune scorpions identifies the Amu Darya River as a long-standing component of Central Asian biogeography. Zoology in the Middle East, 55:95–110.

2011

Bryson, R.B, R.W. Murphy, M.R. Graham, A. Lathrop & D. Lazcano-villarea. 2011. Ephemeral Pleistocene woodlands connect the dots for highland rattlesnakes of the Crotalus intermedius group. Journal of Biogeography, 38:2299–2310.

Graham, M.R. 2011. Book Review – Scorpions of the World. Journal of Arachnology, 39:166–167.

Crother, B.I., M.E. White, J.M. Savage, M.E. Eckstut, M.R. Graham & D.W. Gardner. 2011. A Reevaluation of the Status of the Foxsnakes Pantherophis gloydi Conant and P. vulpinus Baird and Girard (Lepidosauria). ISRN Zoology 1–15. [new species: Pantherophis ramspotti]

2010

Graham, M.R. & R.W. Bryson. 2010. Vaejovis montanus (Scorpiones: Vaejovidae), a new species from the Sierra Madre Occidental of Mexico. Journal of Arachnology, 38:285–293.

Bryson, R.W. & M.R. Graham. 2010. A new alligator lizard from northeastern Mexico. Herpetologica, 66:92–98.

2007

Graham, M.R. & M.E. Soleglad. 2007. A new scorpion genus representing a primitive taxon of tribe Stahnkeini, with a description of a new species from Sonora, Mexico(Scorpiones: Vaejovidae). Euscorpius, 57:1–13. [new genus & species: Gertschius crassicorpus]

Graham, M.R. 2007. Sky Island Vaejovis: two new species and a redescription of V. vorhiesi Stahnke (Scorpiones: Vaejovidae). Euscorpius, 51:1–14. [new species: Vaejovis cashi, V. feti]

2006

Teruel, R., V. Fet & M.R. Graham. 2006. The first mitochondrial DNA phylogeny of Cuban Buthidae (Scorpiones: Buthoidea). Boletín de la S.E.A., 39:219–226.

Graham, M.R. & V. Fet. 2006. Serrula in retrospect: a historical look at scorpion literature (Scorpiones: Orthosterni). Euscorpius, 48:1–19.

Graham, M.R. 2006. Redescription and lectotype designation of Vaejovis lapidicola Stahnke, 1940 (Scorpiones: Vaejovidae). Euscorpius, 46:1–6.

Graham, M.R. 2006. Malformed pedipalp finger dentition of the scorpion Superstitionia donensis (Scorpiones: Superstitioniidae). Euscorpius, 42:1–4.

2005

Fan, J., M.R. Graham, H. Akabane, L.L. Richardson & G. Zhu. 2005. Identification of a novel male germ cell-specific gene TESF-1 in mice. Biochemical and Biophysical Research Communications, 340:8–12.

2003

Fet, E.V., D. Neff, M.R. Graham & V. Fet. 2003. Metasoma of Orthochirus (Scorpiones: Buthidae): are scorpions evolving a new sensory organ? Revista Ibérica de Aracnologia, 8:69–72.